Outbound · the real web of water bears
Sources, not a bibliography cosplay.
Start here when you want a primary page, a community, or a search box. We list Wikipedia, Reddit, YouTube, iNaturalist, taxonomy databases, and literature that actually exists. We do not mint fake DOIs.
Must-visit hubs
- Wikipedia · Tardigradeen.wikipedia.org/wiki/Tardigrade — the best public map of the phylum
- Wikipedia · Tardigrades in spaceFlight list: FOTON-M3, STS-134, Beresheet
- Reddit · r/tardigradesIdentification help, scope photos, community
- iNaturalist · TardigradaObservations, maps, and the reminder that most “tardigrade” photos are moss
Video & search
- YouTube search · tardigradeyoutube.com/results?search_query=tardigrade
- YouTube search · tardigrade microscopeLive walkers beat CGI every time
- YouTube search · Bob Goldstein tardigradeLab footage of Hypsibius development and walking
- YouTube search · FOTON-M3 tardigradeMission context for the 2007 flight
Literature we actually named on this site
- Jönsson et al. 2008 · Current BiologyTardigrades survive exposure to space in low Earth orbit — FOTON-M3 / TARDIS
- Hashimoto et al. 2016 · Nature CommunicationsRamazzottius varieornatus genome and Dsup
- Yoshida et al. 2017 · PLOS BiologyComparative genomics of H. dujardini / H. exemplaris and R. varieornatus
- Gąsiorek et al. 2018 · ZootaxaRedescription and Hypsibius exemplaris
- Boothby et al. 2017 · Molecular CellIntrinsically disordered proteins and desiccation (CAHS)
- Chavez et al. 2019 · eLifeDsup binds nucleosomes and protects DNA from hydroxyl radicals
- Rizzo et al. · TARDIKISSSpace flight effects on antioxidant molecules in dry tardigrades (STS-134)
- Makarova et al. 2026 · Functional & Integrative GenomicsCucumibius annulatus genome and specific proteins
- Kamilari et al. 2026 · Molecular EcologyReview of tardigrade extreme stress tolerance
- Xi et al. 2026 · ACS Applied Materials & InterfacesCAHS motif for glycerol-free red blood cell cryopreservation
- Darch et al. 2026 · Scientific ReportsImproves recovery but not acute stress response of endothelial cells
- Cui et al. 2025 · Frontiers in PharmacologyTardigrade biology and radioprotection drug discovery
- Gąsiorek et al. 2017 · Zoologischer AnzeigerIntegrative redescription of Echiniscus testudo, the nominal taxon for Heterotardigrada
- Jørgensen et al. 2007 · Journal of LimnologyLow DNA sequence diversity in Echiniscus testudo over a large geographical area
- Murai et al. 2021 · BMC GenomicsMultiomics study suggesting possible convergent evolution of abundant heat-soluble proteins
- Roszkowska et al. 2023 · PLOS ONEHow long tardigrades can survive in the anhydrobiotic state
- Erdmann et al. 2021 · PeerJTolerance of Echiniscus testudo and Milnesium inceptum to hypomagnetic conditions
- Weinberg et al. 2025 · BMC GenomicsLineage-specific expansions of the Dicer gene family in tardigrades
- Ito et al. 2025 · bioRxiv (preprint)Preprint: a functional atlas of the tardigrade resistome
- Vecchi & Stec 2025 · Organisms Diversity & EvolutionMitogenome of a new Ramazzottius species from rock pools
- Xi et al. 2025 · bioRxiv (preprint)Preprint: tardigrade CAHS proteins protect synthetic cells under stress
- Guidetti et al. 2016 · Zoological Journal of the Linnean SocietyMoved Richtersius coronifer into a new family, Richtersiidae, within Macrobiotoidea
- Stec & Michalczyk 2020 · ZootaxaNew neotype of R. coronifer from Svalbard; 1981 Bodø neotype invalid under ICZN Art. 75.3.4
- Stec et al. 2020 · Zoological LettersSpitsbergen redescription; seven distinct potential species in Richtersius besides the nominal one
- Pedersen et al. 2021 · AstrobiologyUses Richtersius cf. coronifer (cf. = “compare with”)
- Halberg et al. 2013 · PLOS ONE~87% body-volume reduction into the tun; tun built by muscles
- Jönsson & Rebecchi 2002 · Journal of Experimental ZoologyStorage cells shrank during anhydrobiosis — first evidence energy is used in the process
- Czerneková & Jönsson 2016 · PLOS ONESurvived up to 6 repeated desiccations, with survival falling each cycle
- Jönsson et al. 2005 · International Journal of Radiation Biology0.5 and 1 kGy gamma; authors suggest tolerance may rely on DNA repair
Taxonomy, labs, museums
- WoRMS · TardigradaMarine species database entry for the phylum
- ITIS · TardigradaIntegrated Taxonomic Information System
- NCBI Taxonomy · TardigradaIncluding Hypsibius exemplaris Z151 assemblies
- Encyclopedia of Life · TardigradaAggregated taxon page
- Wikimedia Commons · TardigradaWhere this site’s photographs come from
- ESAFOTON / Biopan mission context
Image licenses are not optional. Every file we ship is listed in images/CREDITS.md with author, license, and source URL. If we cropped a thumbnail, the Commons page is still the source.